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Joseph Gordon-Levitt Tom Zooey Deschanel Summer Geoffrey Arend McKenzie Chloë Grace Moretz Rachel Matthew Gray Gubler Paul Clark Gregg Vance Patricia Belcher Millie Rachel Boston Alison Minka Kelly Autumn Charles Walker Millie's New Husband Ian Reed Kesler Douche Darryl Alan Reed Bus Driver Valente Rodriguez Employee #1 Yvette Nicole Brown New Secretary Nicole Vicius Partygoer Natalie Boren Another Partygoer Maile Flanagan Rhoda Darryl Sivad Usher Gregory A. Thompson Minister Michael Bodie Man John Mackie Mime Jacob Stroop Cupid Kevin Michael Wedding Singer Sid Wilner Grossman Richard McGonagle Narrator (voice) Jean-Paul Vignon French Narrator (voice) Bryan Anthony Dancer Sybil Azur Dancer Cheryl Baxter Dancer Gustavo Carr Dancer John R. Corella Dancer Nadine Ellis Dancer Alejandro Estornel Dancer Nathaniel Flatt Dancer Reshma Gajjar Dancer Tiffany Granath Dancer Jennifer Hamilton Dancer Brandon Henschel Dancer Michael Higgins Dancer Kenneth Hughes Dancer Alexandra Nicole Hulme Dancer John Jacquet Jr. Dancer Jennifer Lee Keyes Dancer Tim Lacatena Dancer Rebecca Lin Dancer Gelsey Weiss Dancer Katie Malia Dancer Anthony Marciona Dancer Christopher Martinez Dancer Vivian Nixon Dancer Tracy Phillips Dancer Nathan Prevost Dancer Jamie Shea Dancer Ryan Thomas Dancer Christian Vincent Dancer Jull Weber Dancer Olivia Howard Bagg Summer - Age 12 (uncredited) Jason Robinson The Puma (uncredited) Chris Connell Marcus (uncredited) Samantha Krutzfeldt Charlie (uncredited) Kathryn Weisbeck Summer Lookalike (uncredited) Eileen Álvarez Summer Lookalike (uncredited) Jennifer Hetrick Sarah (uncredited) Pleasant Wayne Tavern Patron (uncredited)